Here is a list of my publications.


Kristensen T. N., Ketola T. and Kronholm I. Adaptation to environmental stress at different timescales. Annals of the New York Academy of Sciences, online early

Kronholm I. & Ketola T. Effects of acclimation time and epigenetic mechanisms on growth of Neurospora in fluctuating environments. Heredity, 121: 327-341


Richards C. L., Alonso C., Becker C., Bossdorf O., Bucher E., Colomé-Tatché M., Durka W., Engelhardt J., Gaspar B., Gogol-Döring A., Grosse I., van Gurp T. P., Heer K., Kronholm I., Lampei C., Latzel V., Mirouze M., Opgenoorth L., Paun O., Prohaska S., Rensing S. A., Stadler P., Trucchi E., Ullrich K. and Verhoeven K. J. F. 2017. Ecological plant epigenetics: Evidence from model species, and the way forward. Ecology Letters 20: 1576-1590

Kronholm I. 2017. Adaptive evolution and epigenetics. In Tollefsbol T. (ed.), Handbook of Epigenetics: The New Molecular and Medical Genetics (pp. 427-438), 2nd ed., Academic Press, Cambridge. ISBN: 978-0-12-805388-1

Kronholm I., Bassett A., Baulcombe D., and Collins S. 2017. Epigenetic and genetic contributions to adaptation in Chlamydomonas. Molecular Biology and Evolution 34: 2285-2306


Kronholm I., Johannesson H., Ketola T. Epigenetic control of phenotypic plasticity in the filamentous fungus Neurospora crassa. G3: Genes|Genomes|Genetics 6: 4009-4022

Kronholm I. & Collins S. 2016. Epigenetic mutations can both help and hinder adaptive evolution. Molecular Ecology 25: 1856-1868


Saez-Aguayo S., Rondeau-Mouro C., Macquet A., Kronholm I., Ralet M.-C., Berger A., Sallé C., Poulain D., Granier F., Bortan L., Loudet O., de Meaux J., Marion-Poll A. & North H. M. 2014. Local evolution of seed flotation in Arabidopsis. PLoS Genetics 10: e1004221


Kronholm I., Xavier F. P., Alonso-Blanco C., Goudet J. & de Meaux J. Genetic basis of adaptation in Arabidopsis thaliana: local adaptation at the seed dormancy QTL DOG1. 2012. Evolution 66: 2287-2302

Vetter M. M., Kronholm I., He F., Häweker H., Reymond M., Bergelson J., Robatzek S. and de Meaux J. 2012. Flagellin perception varies quantitatively in Arabidopsis thaliana and its relatives. Molecular Biology and Evolution 29: 1655-1667


Chiang G. C. K., Bartsch M., Barua D., Nakabayashi K., Debieu M., Kronholm I., Koornneef M., Soppe W. J. J., Donohue K., and de Meaux J. 2011. DOG1 expression is predicted by the seed-maturation environment and contributes to geographic variation in germination in Arabidopsis thaliana. Molecular Ecology 20: 3336-3349


Alcázar R., García A. V., Kronholm I., de Meaux J., Koornneef M., Parker J. E. and Reymond M. 2010. Natural variation at the Strubbelig Receptor Kinase 3 drives immune-triggered incompatibilities between A. thaliana accessions. Nature Genetics 42: 1135-1139

Kronholm I., Loudet O. and de Meaux J. 2010. Influence of mutation rate on estimators of genetic differentiation – lessons from Arabidopsis thaliana. BMC Genetics 11: 33